Early hominins from Morocco basal to Homo sapiens lineage
Meyer, M. et al. Nuclear DNA sequences from the Middle Pleistocene Sima de los Huesos hominins. Nature 531, 504–507 (2016).
Google Scholar
Duval, M. et al. The first direct ESR dating of a hominin tooth from Atapuerca Gran Dolina TD-6 (Spain) supports the antiquity of Homo antecessor. Quat. Geochronol. 47, 120–137 (2018).
Google Scholar
Bermúdez de Castro, J. M. et al. A hominid from the Lower Pleistocene of Atapuerca, Spain: possible ancestor to Neandertals and modern humans. Science 276, 13921395 (1997).
Google Scholar
Hublin, J.-J. et al. New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens. Nature 546, 289–292 (2017).
Google Scholar
Rightmire, G. P. Human evolution in the Middle Pleistocene: the role of Homo heidelbergensis. Evol. Anthropol. 6, 218–227 (1998).
Google Scholar
Mounier, A. & Caparros, M. The phylogenetic status of Homo heidelbergensis—a cladistic study of Middle Pleistocene hominins. Bull. Mém. Soc. Anthop. Paris 27, 125 (2015).
Bermúdez de Castro, J. M. & Martinón-Torres, M. The origin of the Homo sapiens lineage: when and where? Quat. Int. 634, 1–13 (2022).
Google Scholar
Clarke, R. J. The Ndutu cranium and the origin of Homo sapiens. J. Hum. Evol. 19, 699–736 (1990).
Google Scholar
Texier, J.-P. et al. Lithostratigraphy of the littoral deposits of the last one million years in the Casablanca region (Maroc). Quaternaire 13, 23–41 (2002).
Google Scholar
Lefèvre, D. et al. Les paléolittoraux plio-pléistocènes de Casablanca, cadre chronostratigraphique et paléogéographique de la Préhistoire ancienne du Maroc atlantique. Bull. Archéol. Maroc. 21, 39–70 (2021).
Raynal, J.-P. & Texier, J.-P. Découverte d’Acheuléen ancien dans la carrière Thomas 1 à Casablanca et problème de l’ancienneté de la présence humaine au Maroc. C. R. Acad. Sci. Paris 308, 1743–1749 (1989).
Raynal, J. P., Magoga, L., Sbihi-Alaoui, F.-Z. & Geraads, D. in The Earliest Occupation of Europe Vol. 27: Analecta Praehistorica Leidensia (eds Roebroeks, W. & Van Kolfschoten, T.) Ch. 13, 255–262 (Univ. Leiden, 1995).
Gallotti, R. et al. First high resolution chronostratigraphy for the early North African Acheulean at Casablanca (Morocco). Sci. Rep. 11, 15340 (2021).
Google Scholar
Sausse, F. La mandibule atlanthropienne de la carrière Thomas I (Casablanca). L’Anthropologie 79, 81–112 (1975).
Geraads, D., Beriro, P. & Roche, H. La faune et l’industrie des sites à Homo erectus des Carrières Thomas (Maroc). Précisions sur l’âge de ces Hominidés. C. R. Acad. Sci. Paris 291, 195–198 (1980).
Ennouchi, E. Découverte d’un Pithécanthropien au Maroc. C. R. Acad. Sci. Paris 269, 763–765 (1969).
Raynal, J.-P. et al. Hominid cave at Thomas Quarry I (Casablanca, Morocco): recent findings and their context. Quat. Int. 223-224, 369–382 (2010).
Google Scholar
Raynal, J.-P. et al. in Handbook of Pleistocene Archaeology of Africa: Hominin Behavior, Geography, and Chronology (eds Beyin, A. et al.) 841–861 (Springer, 2023).
Raynal, J. P. et al. Contextes et âge des nouveaux restes dentaires humains du Pléistocène moyen de la carrière Thomas I à Casablanca (Maroc). Bull. Soc. Préhist. Franç. 108, 645–669 (2011).
Google Scholar
Geraads, D. et al. in African Paleoecology and Human Evolution (eds Bobe R. & Reynolds S. C.) 481–491 (Cambridge Univ. Press, 2022).
Daujeard, C. et al. Pleistocene hominins as a resource for carnivores: a c. 500,000-year-old human femur bearing tooth-marks in North Africa (Thomas Quarry I, Morocco). PLoS ONE 11, e0152284 (2016).
Google Scholar
Channell, J. E. T., Singer, B. S. & Jicha, B. R. Timing of Quaternary geomagnetic reversals and excursions in volcanic and sedimentary archives. Quat. Sci. Rev. 228, 106114 (2020).
Google Scholar
Channell, J. E. T. Complexity in Matuyama–Brunhes polarity transitions from North Atlantic IODP/ODP deep-sea sites. Earth Planet. Sci. Lett. 467, 43–56 (2017).
Google Scholar
Haneda, Y., Okada, M., Suganuma, Y. & Kitamura, T. A full sequence of the Matuyama–Brunhes geomagnetic reversal in the Chiba composite section, Central Japan. Prog. Earth Planet. Sci. 7, 44 (2020).
Google Scholar
Lisiecki, L. E. & Raymo, M. E. A Pliocene-Pleistocene stack of 57 globally distributed benthic δ18O records. Paleoceanography 20, PA1003 (2005).
Google Scholar
Pickford, M. The fossil Suidae (Mammalia, Artiodactyla) from Ternifine (Tighenif) Algeria. Münch. Geowiss. Abh. A 50, 1–67 (2020).
Geraads, D. Biochronologie mammalienne du Quaternaire du Maroc atlantique, dans son cadre régional. L’Anthropologie 114, 324–340 (2010).
Google Scholar
Larrasoaña, J. C., Roberts, A. P. & Rohling, E. J. Dynamics of green Sahara periods and their role in hominin evolution. PLoS ONE 8, e76514 (2013).
Google Scholar
Larrasoaña, J. C. A review of West African monsoon penetration during Green Sahara periods; implications for human evolution and dispersals over the last three million years. Oxford Open Climate Change 1, kgab011 (2021).
Google Scholar
Rhodes, E. J., Singarayer, J. S., Raynal, J. P., Westaway, K. E. & Sbihi-Alaoui, F. Z. New age estimates for the Palaeolithic assemblages and Pleistocene succession of Casablanca, Morocco. Quat. Sci. Rev. 25, 2569–2585 (2006).
Google Scholar
Grün, R. & Stringer, C. Direct dating of human fossils and the ever-changing story of human evolution. Quat. Sci. Rev. 322, 108379 (2023).
Google Scholar
Dobson, S. D. & Trinkaus, E. Cross-sectional geometry and morphology of the mandibular symphysis in Middle and Late Pleistocene Homo. J. Hum. Evol. 43, 67–87 (2002).
Google Scholar
Bermúdez de Castro, J. M., Martinón-Torres, M., Gómez-Robles, A. G., Prado, L. & Carbonell, E. New human evidence of the Early Pleistocene settlement of Europe, from Sima del Elefante site (Sierra de Atapuerca, Burgos, Spain). Quat. Int. 223-224, 431–433 (2010).
Google Scholar
Rosas, A. & Bermúdez de Castro, J. M. The ATD6-5 mandibular specimen from Gran Dolina (Atapuerca, Spain). Morphological study and phylogenetic implications. J. Hum. Evol. 37, 567–590 (1999).
Google Scholar
Rosas, A. Occurrence of Neanderthal features in mandibles from the Atapuerca-SH site. Am. J. Phys. Anthropol. 114, 74–91 (2001).
Google Scholar
Vialet, A., Modesto-Mata, M., Martinón-Torres, M., Martínez de Pinillos, M. & Bermúdez de Castro, J. M. A reassessment of the Montmaurin-La Niche mandible (Haute Garonne, France) in the context of European Pleistocene human evolution. PLoS ONE 13, e0189714 (2018).
Google Scholar
Mounier, A., Marchal, F. & Condemi, S. Is Homo heidelbergensis a distinct species? New insight on the Mauer mandible. J. Hum. Evol. 56, 219–246 (2009).
Google Scholar
Rak, Y., Ginzburg, A. & Geffen, E. Does Homo neanderthalensis play a role in modern human ancestry? The mandibular evidence. Am. J. Phys. Anthropol. 119, 199204 (2002).
Google Scholar
Rosas, A. Seventeen new mandibular specimens from the Atapuerca/Ibeas Middle Pleistocene Hominids sample (1985-1992). J. Hum. Evol. 28, 533–559 (1995).
Google Scholar
Rosas, A. & Bermúdez de Castro, J. M. The Mauer mandible and the evolutionary significance of Homo heidelbergensis. Geobios 31, 687–697 (1998).
Google Scholar
Carbonell, E. et al. An Early Pleistocene hominin mandible from Atapuerca-TD6, Spain. Proc. Natl Acad. Sci. USA 102, 5674–5678 (2005).
Google Scholar
Bermúdez de Castro, J. M., Martinón-Torres, M., Gómez-Robles, A., Prado, L. & Sarmiento, S. Comparative analysis of the Gran-Dolina-TD6 (Spain) and Tighennif (Algeria) hominin mandibles. Bull. Mém. Soc. Anthropol. Paris 19, 166–178 (2007).
Google Scholar
Boughner, J. C., Marchiori, D. F. & Packota, G. V. Unexpected variation of human molar size patterns. J. Hum. Evol. 161, 103072 (2021).
Google Scholar
Carretero, J. M., Lorenzo, C. & Arsuaga, J. L. Axial and appendicular skeleton of Homo antecessor. J. Hum. Evol. 37, 459–499 (1999).
Google Scholar
Sahnouni, M. & van der Made, J. in The Cutting Edge: New Approaches to the Archaeology of Human Origins (eds Schick, K. & Toth, N.) 179–210 (Stone Age Institute Press, 2009).
Geraads, D. in African Paleoecology and Human Evolution (eds Bobe, R. & Reynolds, S. C.) 475–480 (Cambridge Univ. Press, 2022).
Geraads, D. Pleistocene Carnivora (Mammalia) from Tighennif (Ternifine), Algeria. Geobios 49, 445–458 (2016).
Google Scholar
Martini, P. & Geraads, D. Camelus thomasi Pomel, 1893 from the Pleistocene typelocality Tighennif (Algeria). Comparisons with modern Camelus. Geodiversitas 40, 115–134 (2018).
Google Scholar
Martinón-Torres, M. et al. in Dental Perspectives on Human Evolution: State of the Art Research in Dental Paleoanthropology (eds Bailey, S. & Hublin, J.-J.) Ch. 5, 65–79 (Springer, 2007).
Bermúdez de Castro, J. M. et al. Comparative dental study between Homo antecessor and Chinese Homo erectus: Nonmetric features and geometric morphometrics. J. Hum. Evol. 161, 103087 (2021).
Google Scholar
Gómez-Robles, A. Dental evolutionary rates and its implications for the Neanderthal modern human divergence. Sci. Adv. 5, eaaw1268 (2019).
Google Scholar
Feng, X. et al. The phylogenetic position of the Yunxian cranium elucidates the origin of Homo longi and the Denisovans. Science 389, 1320–1324 (2025).
Google Scholar
Harrison, R. J., & Feinberg, J. M. FORCinel: an improved algorithm for calculating first-order reversal curve distributions using locally weighted regression smoothing. Geochem. Geophys. Geosyst. https://doi.org/10.1029/2008GC001987 (2008).
Bergmann, I., Hublin, J.-J., Gunz, P. & Freidline, S. E. How did modern morphology evolve in the human mandible? The relationship between static adult allometry and mandibular variability in Homo sapiens. J. Hum. Evol. 157, 103026 (2021).
Google Scholar
Bergmann, I. et al. The relevance of late MSA mandibles on the emergence of modern morphology in Northern Africa. Sci. Rep. 12, 8841 (2022).
Google Scholar
Freidline, S. E., Gunz, P., Harvati, K. & Hublin, J.-J. Middle Pleistocene human facial morphology in an evolutionary and developmental context. J. Hum. Evol. 63, 723–740 (2012).
Google Scholar
Freidline, S. E., Gunz, P., Harvati, K. & Hublin, J.-J. Evaluating developmental shape changes in Homo antecessor subadult facial morphology. J. Hum. Evol. 65, 404–423 (2013).
Google Scholar
Freidline, S. E., Gunz, P., Janković, I., Harvati, K. & Hublin, J. J. A comprehensive morphometric analysis of the frontal and zygomatic bone of the Zuttiyeh fossil from Israel. J. Hum. Evol. 62, 225–241 (2012).
Google Scholar
Freidline, S. E. et al. Early presence of Homo sapiens in Southeast Asia by 86-68 kyr at Tam Pa Ling, Northern Laos. Nat. Commun. 14, 3193 (2023).
Google Scholar
Wiley, D. F. et al. In Proc. 16th IEEE Visualization Conference (VIS 2005) Vol. 55, 431–438 (IEEE, 2005).
R Core Team. R: A Language and Environment for Statistical Computing (R Foundation for Statistical Computing, 2021); www.R-project.org/.
Schlager, S. in Statistical Shape and Deformation Analysis (eds Zheng, S. L. G. & Szekely, G.) 217–256 (Academic, 2017).
Adams, D. et al. Geomorph: software for geometric morphometric analyses. R package version 4.0.2 (2021); cran.rproject.org/package=geomorph.
Baken, E. K. et al. geomorph v4.0 and gmShiny: enhanced analytics and a new graphical interface for a comprehensive morphometric experience. Methods Ecol. Evol. 12, 2355–2363 (2021).
Google Scholar
Gunz, P. et al. Principles for the virtual reconstruction of hominin crania. J. Hum. Evol. 57, 48–62 (2009).
Google Scholar
Gunz, P., Mitteroecker P. & Bookstein F. in Modern Morphometrics in Physical Anthropology (ed. Slice, D. E.) 73–98 (Plenum, 2005).
Gunz, P. & Mitteroecker, P. Semilandmarks: a method for quantifying curves and surfaces. Hystrix 24, 103–109 (2013).
Rohlf, F. J. & Slice, D. Extensions of the Procrustes method for the optimal superimposition of landmarks. Syst. Biol. 39, 40–59 (1990).
Arambourg, C. & Biberson, P. The fossil human remains from the Paleolithic site of Sidi Abderrahman (Morocco). Am. J. Phys. Anthropol. 14, 467–489 (1956).
Google Scholar
Rightmire, G. P. Comparative studies of Late Pleistocene human remains from Klasies River Mouth, South Africa. J. Hum. Evol. 20, 131–156 (1991).
Google Scholar
Stewart, T. D. in Annual Report of the Smithsonian Institution 521–533 (US Government Printing Office, 1962).
Tillier, A.-M. in Le Squelette Moustérien de Kébara 2 97–112 (CNRS, 1991).
Stewart, T. D. The Neanderthal skeletal remains from Shanidar Cave, Iraq: a summary of findings to date. Proc. Am. Phil. Soc. 121, 121–165 (1977).
Trinkaus, E. The Shanidar Neanderthals (Academic, 1983).
Leroi-Gourhan, A. Étude des Restes Humains Fossiles provenant des Grottes d’Arcysur-Cure (Masson et Cie, 1958).
Daura, J. et al. A Neandertal mandible from the Cova del Gegant (Sitges, Barcelona, Spain). J. Hum. Evol. 49, 56–70 (2005).
Google Scholar
Topinard, P. Les caractères simiens de la mâchoire de la Naulette. Rev. Antropol. 15, 385–431 (1886).
Blake, C. C. On a human jaw from the cave of La Naulette, near Dinant, Belgium. Anthropol. Rev. 5, 294–303 (1867).
Google Scholar
Leguebe, A. & Toussaint, M. in La Mandibule et les Cubitus de la Naulette: Morphologie et Morphométrie 15 (CNRS, 1988).
Heim, J. L. Les hommes fossiles de La Ferrassie (Dordogne) et le problème de la définition des Neandertaliens classiques. III. Squelette céphalique. L’Anthropologie 78, 321–378 (1974).
de Lumley, M.-A. Les Néandertaliens de la grotte de l’Hortus. Etud. Quat. 1, 375–385 (1972).
Condemi, S. et al. Possible interbreeding in late Italian Neanderthals? New data from the Mezzena jaw (Monti Lessini, Verona, Italy). PLoS ONE 8, e59781 (2013).
Google Scholar
Corrain, C. Resti scheletrici umani del ‘Riparo Mezzena’. Mem. Mus. Civ. Stor. Nat. Verona 16, 97–101 (1968).
Walker, M. J., Lombardi, A. V., Zapata, J. & Trinkaus, E. Neandertal mandibles from the Sima de las Palomas del Cabezo Gordo, Murcia, southeastern Spain. Am. J. Phys. Anthropol. 142, 261–272 (2010).
Google Scholar
Martin, H. Machoire humaine moustérienne trouvée dans la station de La Quina. L’homme Préhist. 13, 3–21 (1926).
Martin, H. Position stratigraphique des Ossements humains recueillis dans le Moustérien de La Quina de 1908 à 1912. Bull. Soc. Préhist. Fr. 9, 700–709 (1912).
Martin, H. L.’Homme Fossile de la Quina (Libraire Octave Doin, 1923).
Pap, I., Tillier, A. M., Arensburg, B. & Chech, M. The Subalyuk Neanderthal remains (Hungary): a re-examination. Ann. Hist. Nat. Mus. Natl Hung. 88, 233–270 (1996).
Sanchez, F. Comparative biometrical study of the Mousterian mandible from Cueva del Boquete de Zafarraya (Málaga, Spain). J. Hum. Evol. 14, 125–138 (1999).
Google Scholar
Vlček, E. Fossile Menschenfunde von Weimar-Ehringsdorf, Weimarer Monographien zur Ur- und Frühgeschichte Vol. 30 (Landesamt für Archäologische Denkmalpflege, 1993).
Condemi, S. Les Néandertaliens de La Chaise: Abri Bourgeois-Delaunay (CTHS, 2001).
Bar-Yosef, O. & Vandermeersch, B. (eds) Le Squelette Moustérien de Kébara 2 (CNRS, 1991).
Arambourg, C. & Hoffstetter, R. Le Gisement de Ternifine Vol. 1 (Masson, 1963).
Hooton, E. A., Hencken, H. O. & Snow, C.hE. The ancient Palestinian: Skhūl V reconstruction. Am. School Prehist. Res. 17, 5–10 (1953).
Sollas, W. J. The Chancelade skull. J. R. Anthropol. Inst. 57, 89–122 (1927).
Martin, H. Caractères des squelettes humains quaternaires de la vallée du Roc (Charente). Bull. Mem. Soc. Anthropol. Paris 8, 103–129 (1927).
Google Scholar
Vercellotti, G., Alciati, G., Richards, M. P. & Formicola, V. The Late Upper Paleolithic skeleton Villabruna 1 (Italy): a source of data on biology and behavior of a 14,000year-old hunter. J. Anthropol. Sci. 86, 143–163 (2008).
Google Scholar
Formicola, V. Una mandibola umana dal deposito dell’Epigravettiano finale delle Arene Candite (scavi 1970). Rev. Antropol. 64, 271–278 (1986).
Odano, A. M. & Riquet, R. Le gisement préhistorique de Dar-es-Soltane 2. Champ de tir de El Menzeh à Rabat (Maroc). Note préliminaire. 2-Étude anthropologique des restes post-atériens. Bull. d’Archéologie Maroc. 11, 25–63 (1978).
Debénath, A. Nouveaux restes humains atériens du Maroc. C. R. Acad. Sci., Paris 290, 851–852 (1980).
Google Scholar
Crognier, E. & Dupouy-Madre, M. Les Natoufiens du Nahal Oren (Ouadi Fallah). Etude anthropologique. Paléorient 2, 103–121 (1974).
Google Scholar
Henke, W. Vergleichend-morphologische Kennzeichnung der Jungpaläolithiker von Oberkassel bei Bonn. Z. Morphol. Anthropol. 75, 27–44 (1984).
Google Scholar
Hershkovitz, I. et al. Ohalo II H2: a 19,000-year-old skeleton from a water-logged site at the Sea of Galilee, Israel. Am. J. Phys. Anthropol. 96, 215–234 (1995).
Google Scholar
Soficaru, A., Dobos, A. & Trinkaus, E. Early modern humans from the Pestera Muierii, Baia de Fier, Romania. Proc. Natl Acad. Sci. USA 103, 17196–17201 (2006).
Google Scholar
Crevecoeur, I. Etude Anthropologique des Restes Humains de Nazlet Khater (Paléolithique Superieur, Egypte). PhD Thesis, Université Sciences et Technologies Bordeaux I (2006).
Thoma, A. Morphology and affinities of the Nazlet Khater man. J. Hum. Evol. 13, 287–296 (1984).
Google Scholar
Anderson, J. E. in The Prehistory of Nubia Vol. 2 (ed. Wendorf, F.) 996–1040 (Southern Methodist Univ. Press, 1968).
Crevecoeur, I. From the Nile to the Danube: a comparison of the Nazlet Khater 2 and Oase 1 early modern human mandibles. Anthropologie 42, 203–213 (2004).
Trinkaus, E. et al. An early modern human from the Pestera cu Oase, Romania. Proc. Natl Acad. Sci. USA 100, 11231–11236 (2003).
Google Scholar
Trinkaus, E. & Svoboda, J. (eds) Early Modern Human Evolution in Central Europe. The People of Dolní Věstonice and Pavlov. The Dolni Vestonice Studies Vol. 12 (Oxford Univ. Press, 2006).
Sládek, V., Trinkaus, E., Hillson, S. W. & Holliday, T. W. The People of the Pavlovian. Skeletal Catalogue and Osteometrics of the Gravettian Fossil Hominids from Dolní Věstonice and Pavlov. The Dolni Vestonice Studies Vol. 5 1–244 (Brno, 2000).
Drozdová, E. The evaluation of a rediscovered fragment of human lower jaw, No 21 from Předmostí u Přerova. Archeol. Rozhl. 53, 452–460 (2001).
Dutour, O. Palimpseste paléoanthropologique sur l’‘Homme fossile d’Asselar’ (Sahara). Préhist. Anthropol. Médit. 1, 73–83 (1992).
Gambier, D. Vestiges humains du gisement d’Isturitz (Pyrénées-Atlantiques): étude anthropologique et analyse des traces d’action humaine intentionnelle. Antiqu. Natl. 22–23, 9–26 (1990).
Wood, B. Koobi Fora Research Project, Vol. 4: Hominid Crania Remains (Clarendon, 1991).
Rightmire, G. P. The Evolution of Homo Erectus: Comparative Anatomical Studies of an Extinct Human Species (Cambridge Univ. Press, 1990).
Wollny, G. et al. MIA—a free and open-source software for gray scale medical image analysis. Source Code Biol. Med. 8, 20 (2013).
Google Scholar
Cannoodt, R. & Bengtsson, H. rcannood/princurve: princurve 2.1.4. Zenodo https://doi.org/10.5281/zenodo.3351282 (2019).
Davies, T. W. tomdavies15/EDJ_GM: geometric morphometric analysis of enameldentine junction morphology. Zenodo https://doi.org/10.5281/zenodo.10255289 (2023).
Dryden, I. & Mardia, K. V. Statistical Shape Analysis (Wiley, 1998).
Molnar, S. Human tooth wear, tooth function and cultural variability. Am. J. Phys. Anthropol. 34, 175–189 (1971).
Google Scholar
Turner, C., Nichol, C. & Scott, G. in Advances in Dental Anthropology (eds Kelley, M. & Larsen, C.) 13–31 (Wiley Liss, 1991).
Skinner, M. M. et al. Dental trait expression at the enamel-dentine junction of lower molars in extant and fossil hominoids. J. Hum. Evol. 54, 173–186 (2008).
Google Scholar
Bailey, S., Skinner, M. M. & Hublin, J. J. What lies beneath? An evaluation of lower molar trigonid crest patterns based on both dentine and enamel expression. Am. J. Phys. Anthropol. 145, 505–518 (2011).
Google Scholar
Bailey, S. E. Neandertal Dental Morphology: Implications for Modern Human Origins. PhD thesis, Arizona State University (2002).
Burnett, S. E., Hawkey, D. E. & Turner, C. G. II Population variation in human maxillary premolar accessory ridges (MxPAR). Am. J. Phys. Anthropol. 141, 319–324 (2010).
Google Scholar
Benazzi, S. et al. Early dispersal of modern humans in Europe and implications for Neanderthal behaviour. Nature 479, 525–528 (2011).
Google Scholar
Benazzi, S. et al. Comparison of dental measurement systems for taxonomic assignment of Neanderthal and modern human lower second deciduous molars. J. Hum. Evol. 61, 320–326 (2011b).
Google Scholar
Benazzi, S. et al. Cervical and crown outline analysis of worn Neanderthal and modern human lower second deciduous molars. Am. J. Phys. Anthropol. 149, 537–546 (2012).
Google Scholar
Bailey, S. E., Benazzi, S., Buti, L. & Hublin, J. J. Allometry, merism, and tooth shape of the lower second deciduous molar and first permanent molar. Am. J. Phys. Anthropol. 159, 93–105 (2016).
Google Scholar
Bailey, S. E., Sorrentino, R., Mancuso, G., Hublin, J. J. & Benazzi, S. Taxonomic differences in deciduous lower first molar crown outlines of Homo sapiens and Homo neanderthalensis. J. Hum. Evol. 147, 102864 (2020).
Google Scholar
Jolliffe, I. T. Principal Component Analysis https://doi.org/10.1007/b98835 (Springer, 2002).
Le Cabec, A., Gunz, P., Kupczik, K., Braga, J. & Hublin, J.-J. Anterior tooth root morphology and size in Neanderthals: taxonomic and functional implications. J. Hum. Evol. 64, 169–193 (2013).
Google Scholar
Meyer, M. R. & Haeusler, M. Spinal cord evolution in early Homo. J. Hum. Evol. 88, 43–53 (2015).
Google Scholar
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